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C)$l};) -= KI'Å}Jh;KMŗҫo///./sŗ3KIp!߻,\3n/A+]Vt %Kߨ\**bwF+{V[vH-OK-K-륖R˕RrerR)I-RKTjyJ:M"E᭍fI-{'fTj)ZT{'0Aڇ=y_WC)4#g0-l3O^ >nx gF g:`& 0 QՏkP>tQXU$*ƕ*>Pg[rh~ /Pܜ' qF\CD :o|\C]M,[,zs7ڶҧCӥ_P ΓjT*=9tzx$k;;mF[uygp>B@e{B %<`h/48@G#w:Ҝ!g:NSwNIδ@Ёiz _"U2_ſ,2TeΞ))q2Zh'Q ODr(ODɓrD)]Q(Ƣ\+JR8yf;TПRM U4l.0jm9;Ys[6.jltM|{PQfj6i܆AAÊjhl1e]+cd63e9=˪j,>~f'7dvX8A=QT6z[}:a CJAߺu;G[rEQְ _3~"npZ|=y)ј֊cZ&L@,~֓0gX 8PQb9l0\Vģ"£E~]HKQi-3ID44}V+RlaUL?3XzbE͔Hl7I2bKE9+ MQ*U4s@OVҰ"ׄ]Κv a;J?7f endstream endobj 3 0 obj <> endobj 1 0 obj <> endobj 4 0 obj <>stream uuid:3fc3727c-26c7-495e-9c86-20f5e7f19a8e uuid:C82794B7-9BA8-C7D9-E229-2C049FE6EFA5 True body size; genome size; metabolic rate; repetitive elements; Rotifera PDFlib+PDI 9.0.7p3 (C++/Win64); modified using iTextSharp 4.1.6 by 1T3XT application/pdf 10.1093/gbe/evaf041 en Oxford University Press Comparing Small and Large Genomes Within Monogonont Rotifers Jonathon E. Mohl Patrick D. Brown Aaron J. Robbins Philip Lavretsky Rick Hochberg Robert L. Wallace Elizabeth J. Walsh DOI: 10.1093/gbe/evaf041; Genome Biology and Evolution, 17, 3, 2025-03-06.; Abstract: Genome size is an important correlate of many biological features including body size, metabolic rate, and developmental rate and can vary due to a variety of mechanisms, including incorporation of repetitive elements, duplication events, or reduction due to selective constraints. Our ability to understand the causes of genome size variation is hampered by limited sampling of many nonmodel taxa, including monogonont rotifers. Here, we used high-throughput Nanopore sequencing and flow cytometry to estimate genome sizes of nine species of monogonont rotifers representing seven families, including three representatives of Superorder Gnesiotrocha. We annotated the genomes and classified the repetitive elements. We also compared genome size with two biological features: body size and metabolic rate. Body sizes were obtained from the literature and our estimates. Oxygen consumption was used as a proxy for metabolic rate and was determined using a respirometer. We obtained similar genome size estimates from genome assemblies and flow cytometry, which were positively correlated with body size and size-specific respiration rate. Importantly, we determined that genome size variation is not due to increased numbers of repetitive elements or large regions of duplication. Instead, we observed higher numbers of predicted proteins as genome size increased, but currently many have no known function. Our results substantially expand the taxonomic scope of available genomes for Rotifera and provide opportunities for addressing genetic mechanisms underlying evolutionary and ecological processes in the phylum. © The Author(s) 2025. Published by Oxford University Press on behalf of Society for Molecular Biology and Evolution. body size genome size metabolic rate repetitive elements Rotifera Journal Genome Biology and Evolution © The Author(s) 2025. 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<>/Border[0 0 0]>> endobj 626 0 obj <> endobj 418 0 obj <>/Border[0 0 0]>> endobj 419 0 obj <>/Border[0 0 0]>> endobj 420 0 obj <>/Border[0 0 0]>> endobj 627 0 obj <> endobj 421 0 obj <>/Border[0 0 0]>> endobj 628 0 obj <> endobj 422 0 obj <>/Border[0 0 0]>> endobj 423 0 obj <>/Border[0 0 0]>> endobj 424 0 obj <>/Border[0 0 0]>> endobj 629 0 obj <> endobj 425 0 obj <>/Border[0 0 0]>> endobj 426 0 obj <>/Border[0 0 0]>> endobj 630 0 obj <> endobj 427 0 obj <>/Border[0 0 0]>> endobj 428 0 obj <>/Border[0 0 0]>> endobj 631 0 obj <> endobj 429 0 obj <>/Border[0 0 0]>> endobj 430 0 obj <>/Border[0 0 0]>> endobj 632 0 obj <> endobj 431 0 obj <>/Border[0 0 0]>> endobj 633 0 obj <> endobj 432 0 obj <>/Border[0 0 0]>> endobj 433 0 obj <>/Border[0 0 0]>> endobj 634 0 obj <> endobj 434 0 obj <>/Border[0 0 0]>> endobj 435 0 obj <>/Border[0 0 0]>> endobj 635 0 obj <> endobj 436 0 obj <>/Border[0 0 0]>> endobj 636 0 obj <> endobj 437 0 obj <>/Border[0 0 0]>> endobj 438 0 obj <>/Border[0 0 0]>> endobj 637 0 obj <> endobj 439 0 obj <>/Border[0 0 0]>> endobj 440 0 obj <>/Border[0 0 0]>> endobj 638 0 obj <> endobj 441 0 obj <>/Border[0 0 0]>> endobj 442 0 obj <>/Border[0 0 0]>> endobj 443 0 obj <>/Border[0 0 0]>> endobj 639 0 obj <> endobj 444 0 obj <>/Border[0 0 0]>> endobj 445 0 obj <>/Border[0 0 0]>> endobj 640 0 obj <> endobj 446 0 obj <>/Border[0 0 0]>> endobj 447 0 obj <>/Border[0 0 0]>> endobj 641 0 obj <> endobj 448 0 obj <>/Border[0 0 0]>> endobj 449 0 obj <>/Border[0 0 0]>> endobj 450 0 obj <>/Border[0 0 0]>> endobj 642 0 obj <> endobj 451 0 obj <>/Border[0 0 0]>> endobj 643 0 obj <> endobj 452 0 obj <>/Border[0 0 0]>> endobj 453 0 obj <>/Border[0 0 0]>> endobj 644 0 obj <> endobj 454 0 obj <>/Border[0 0 0]>> endobj 645 0 obj <> endobj 455 0 obj <>/Border[0 0 0]>> endobj 456 0 obj <>/Border[0 0 0]>> endobj 646 0 obj <> endobj 457 0 obj <>/Border[0 0 0]>> endobj 458 0 obj <>/Border[0 0 0]>> endobj 647 0 obj <> endobj 459 0 obj <>/Border[0 0 0]>> endobj 648 0 obj <> endobj 460 0 obj <>/Border[0 0 0]>> endobj 461 0 obj <>/Border[0 0 0]>> endobj 462 0 obj <>/Border[0 0 0]>> endobj 649 0 obj <> endobj 463 0 obj <>/Border[0 0 0]>> endobj 464 0 obj <>/Border[0 0 0]>> endobj 650 0 obj <> endobj 465 0 obj <>/Border[0 0 0]>> endobj 651 0 obj <> endobj 466 0 obj <>/Border[0 0 0]>> endobj 467 0 obj <>/Border[0 0 0]>> endobj 468 0 obj <>/Border[0 0 0]>> endobj 652 0 obj <> endobj 469 0 obj <>/Border[0 0 0]>> endobj 653 0 obj <> endobj 470 0 obj <>/Border[0 0 0]>> endobj 471 0 obj <>/Border[0 0 0]>> endobj 654 0 obj <> endobj 472 0 obj <>/Border[0 0 0]>> endobj 655 0 obj <> endobj 473 0 obj <>/Border[0 0 0]>> endobj 474 0 obj <>/Border[0 0 0]>> endobj 475 0 obj <>/Border[0 0 0]>> endobj 656 0 obj <> endobj 476 0 obj <>/Border[0 0 0]>> endobj 477 0 obj <>/Border[0 0 0]>> endobj 657 0 obj <> endobj 478 0 obj <>/Border[0 0 0]>> endobj 479 0 obj <>/Border[0 0 0]>> endobj 658 0 obj <> endobj 480 0 obj <>/Border[0 0 0]>> endobj 659 0 obj <> endobj 481 0 obj <>/Border[0 0 0]>> endobj 482 0 obj <>/Border[0 0 0]>> endobj 483 0 obj <>/Border[0 0 0]>> endobj 484 0 obj <>/Border[0 0 0]>> endobj 660 0 obj <> endobj 485 0 obj <>/Border[0 0 0]>> endobj 486 0 obj <>/Border[0 0 0]>> endobj 661 0 obj <> endobj 487 0 obj <>/Border[0 0 0]>> endobj 662 0 obj <> endobj 488 0 obj <>/Border[0 0 0]>> endobj 489 0 obj <>/Border[0 0 0]>> endobj 663 0 obj <> endobj 490 0 obj <>/Border[0 0 0]>> endobj 491 0 obj <>/Border[0 0 0]>> endobj 664 0 obj <> endobj 492 0 obj <>/Border[0 0 0]>> endobj 665 0 obj <> endobj 493 0 obj <>/Border[0 0 0]>> endobj 494 0 obj <>/Border[0 0 0]>> endobj 495 0 obj <>/Border[0 0 0]>> endobj 666 0 obj <> endobj 496 0 obj <>/Border[0 0 0]>> endobj 497 0 obj <>/Border[0 0 0]>> endobj 667 0 obj <> endobj 502 0 obj <>/Border[0 0 0]>> endobj 503 0 obj <>/Border[0 0 0]>> endobj 668 0 obj <> endobj 504 0 obj <>/Border[0 0 0]>> endobj 669 0 obj <> endobj 505 0 obj <>/Border[0 0 0]>> endobj 506 0 obj <>/Border[0 0 0]>> endobj 670 0 obj <> endobj 507 0 obj <>/Border[0 0 0]>> endobj 671 0 obj <> endobj 508 0 obj <>/Border[0 0 0]>> endobj 509 0 obj <>/Border[0 0 0]>> endobj 672 0 obj <> endobj 510 0 obj <>/Border[0 0 0]>> endobj 511 0 obj <>/Border[0 0 0]>> endobj 673 0 obj <> endobj 512 0 obj <>/Border[0 0 0]>> endobj 674 0 obj <> endobj 513 0 obj <>/Border[0 0 0]>> endobj 514 0 obj <>/Border[0 0 0]>> endobj 675 0 obj <> endobj 515 0 obj <>/Border[0 0 0]>> endobj 676 0 obj <> endobj 516 0 obj <>/Border[0 0 0]>> endobj 517 0 obj <>/Border[0 0 0]>> endobj 677 0 obj <> endobj 518 0 obj <>/Border[0 0 0]>> endobj 678 0 obj <> endobj 519 0 obj <>/Border[0 0 0]>> endobj 520 0 obj <>/Border[0 0 0]>> endobj 679 0 obj <> endobj 521 0 obj <>/Border[0 0 0]>> endobj 522 0 obj <>/Border[0 0 0]>> endobj 680 0 obj <> endobj 523 0 obj <>/Border[0 0 0]>> endobj 681 0 obj <> endobj 524 0 obj <>/Border[0 0 0]>> endobj 525 0 obj <>/Border[0 0 0]>> endobj 682 0 obj <> endobj 526 0 obj <>/Border[0 0 0]>> endobj 527 0 obj <>/Border[0 0 0]>> endobj 683 0 obj <> endobj 528 0 obj <>/Border[0 0 0]>> endobj 684 0 obj <> endobj 529 0 obj <>/Border[0 0 0]>> endobj 530 0 obj <>/Border[0 0 0]>> endobj 685 0 obj <> endobj 531 0 obj <>/Border[0 0 0]>> endobj 532 0 obj <>/Border[0 0 0]>> endobj 686 0 obj <> endobj 533 0 obj <>/Border[0 0 0]>> endobj 534 0 obj <>/Border[0 0 0]>> endobj 687 0 obj <> endobj 535 0 obj <>/Border[0 0 0]>> endobj 536 0 obj <>/Border[0 0 0]>> endobj 537 0 obj <>/Border[0 0 0]>> endobj 688 0 obj <> endobj 538 0 obj <>/Border[0 0 0]>> endobj 539 0 obj <>/Border[0 0 0]>> endobj 689 0 obj <> endobj 540 0 obj <>/Border[0 0 0]>> endobj 541 0 obj <>/Border[0 0 0]>> endobj 690 0 obj <> endobj 691 0 obj <> endobj 43 0 obj <> endobj 284 0 obj <> endobj 693 0 obj <> endobj 694 0 obj <> endobj 702 0 obj <> endobj 703 0 obj <> endobj 704 0 obj <> endobj 705 0 obj <> endobj 706 0 obj <> endobj 707 0 obj <> endobj 708 0 obj <> endobj 709 0 obj <> endobj 710 0 obj <> endobj 711 0 obj <> endobj 712 0 obj <> endobj 713 0 obj <> endobj 714 0 obj <> endobj 715 0 obj <> endobj 716 0 obj <> endobj 717 0 obj <> endobj 718 0 obj <> endobj 719 0 obj <> endobj 720 0 obj <> endobj 721 0 obj <> endobj 722 0 obj <> endobj 723 0 obj <> endobj 724 0 obj <> endobj 725 0 obj <> endobj 726 0 obj <> endobj 727 0 obj <> endobj 728 0 obj <> endobj 729 0 obj <> endobj 730 0 obj <> endobj 731 0 obj <> endobj 732 0 obj <> endobj 733 0 obj <> endobj 695 0 obj <> endobj 734 0 obj <> endobj 735 0 obj <> endobj 736 0 obj <> endobj 737 0 obj <> endobj 738 0 obj <> endobj 739 0 obj <> endobj 740 0 obj <> endobj 741 0 obj <> endobj 742 0 obj <> endobj 743 0 obj <> endobj 744 0 obj <> endobj 745 0 obj <> endobj 746 0 obj <> endobj 747 0 obj <> endobj 748 0 obj <> endobj 749 0 obj <> endobj 750 0 obj <> endobj 751 0 obj <> endobj 752 0 obj <> endobj 753 0 obj <> endobj 754 0 obj <> endobj 755 0 obj <> endobj 756 0 obj <> endobj 757 0 obj <> endobj 758 0 obj <> endobj 759 0 obj <> endobj 760 0 obj <> endobj 761 0 obj <> endobj 762 0 obj <> endobj 763 0 obj <> endobj 764 0 obj <> endobj 765 0 obj <> endobj 696 0 obj <> endobj 766 0 obj <> endobj 767 0 obj <> endobj 768 0 obj <> endobj 769 0 obj <> endobj 770 0 obj <> endobj 771 0 obj <> endobj 772 0 obj <> endobj 773 0 obj <> endobj 774 0 obj <> endobj 775 0 obj <> endobj 776 0 obj <> endobj 777 0 obj <> endobj 778 0 obj <> endobj 779 0 obj <> endobj 780 0 obj <> endobj 781 0 obj <> endobj 782 0 obj <> endobj 783 0 obj <> endobj 784 0 obj <> endobj 785 0 obj <> endobj 786 0 obj <> endobj 787 0 obj <> endobj 788 0 obj <> endobj 789 0 obj <> endobj 790 0 obj <> endobj 791 0 obj <> endobj 792 0 obj <> endobj 793 0 obj <> endobj 794 0 obj <> endobj 795 0 obj <> endobj 796 0 obj <> endobj 797 0 obj <> endobj 697 0 obj <> endobj 798 0 obj <> endobj 799 0 obj <> endobj 800 0 obj <> endobj 801 0 obj <> endobj 802 0 obj <> endobj 803 0 obj <> endobj 804 0 obj <> endobj 805 0 obj <> endobj 806 0 obj <> endobj 807 0 obj <> endobj 808 0 obj <> endobj 809 0 obj <> endobj 810 0 obj <> endobj 811 0 obj <> endobj 812 0 obj <> endobj 813 0 obj <> endobj 814 0 obj <> endobj 815 0 obj <> endobj 816 0 obj <> endobj 817 0 obj <> endobj 818 0 obj <> endobj 819 0 obj <> endobj 820 0 obj <> endobj 821 0 obj <> endobj 822 0 obj <> endobj 823 0 obj <> endobj 824 0 obj <> endobj 825 0 obj <> endobj 826 0 obj <> endobj 827 0 obj <> endobj 828 0 obj <> endobj 829 0 obj <> endobj 698 0 obj <> endobj 830 0 obj <> endobj 831 0 obj <> endobj 832 0 obj <> endobj 833 0 obj <> endobj 834 0 obj <> endobj 835 0 obj <> endobj 836 0 obj <> endobj 837 0 obj <> endobj 838 0 obj <> endobj 839 0 obj <> endobj 840 0 obj <> endobj 841 0 obj <> endobj 842 0 obj <> endobj 843 0 obj <> endobj 844 0 obj <> endobj 845 0 obj <> endobj 846 0 obj <> endobj 847 0 obj <> endobj 848 0 obj <> endobj 849 0 obj <> endobj 850 0 obj <> endobj 851 0 obj <> endobj 852 0 obj <> endobj 853 0 obj <> endobj 854 0 obj <> endobj 855 0 obj <> endobj 856 0 obj <> endobj 857 0 obj <> endobj 858 0 obj <> endobj 859 0 obj <> endobj 860 0 obj <> endobj 861 0 obj <> endobj 699 0 obj <> endobj 862 0 obj <> endobj 863 0 obj <> endobj 864 0 obj <> endobj 865 0 obj <> endobj 866 0 obj <> endobj 867 0 obj <> endobj 868 0 obj <> endobj 869 0 obj <> endobj 870 0 obj <> endobj 871 0 obj <> endobj 872 0 obj <> endobj 873 0 obj <> endobj 874 0 obj <> endobj 875 0 obj <> endobj 876 0 obj <> endobj 877 0 obj <> endobj 878 0 obj <> endobj 879 0 obj <> endobj 880 0 obj <> endobj 881 0 obj <> endobj 882 0 obj <> endobj 883 0 obj <> endobj 884 0 obj <> endobj 885 0 obj <> endobj 886 0 obj <> endobj 887 0 obj <> endobj 888 0 obj <> endobj 889 0 obj <> endobj 890 0 obj <> endobj 891 0 obj <> endobj 892 0 obj <> endobj 893 0 obj <> endobj 700 0 obj <> endobj 894 0 obj <> endobj 895 0 obj <> endobj 896 0 obj <> endobj 897 0 obj <> endobj 898 0 obj <> endobj 899 0 obj <> endobj 900 0 obj <> endobj 901 0 obj <> endobj 902 0 obj <> endobj 903 0 obj <> endobj 904 0 obj <> endobj 905 0 obj <> endobj 906 0 obj <> endobj 907 0 obj <> endobj 908 0 obj <> endobj 909 0 obj <> endobj 910 0 obj <> endobj 911 0 obj <> endobj 912 0 obj <> endobj 913 0 obj <> endobj 914 0 obj <> endobj 915 0 obj <> endobj 916 0 obj <> endobj 917 0 obj <> endobj 918 0 obj <> endobj 919 0 obj <> endobj 920 0 obj <> endobj 921 0 obj <> endobj 922 0 obj <> endobj 923 0 obj <> endobj 924 0 obj <> endobj 925 0 obj <> endobj 701 0 obj <> endobj 926 0 obj <> endobj 927 0 obj <> endobj 928 0 obj <> endobj 929 0 obj <> endobj 930 0 obj <> endobj 931 0 obj <> endobj 932 0 obj <> endobj 933 0 obj <> endobj 934 0 obj <> endobj 935 0 obj <> endobj 936 0 obj <> endobj 937 0 obj <> endobj 938 0 obj <> endobj 939 0 obj <> endobj 940 0 obj <> endobj 941 0 obj <> endobj 942 0 obj <> endobj 943 0 obj <> endobj 944 0 obj <> endobj 946 0 obj <> endobj 947 0 obj <> 2]>> endobj 1323 0 obj <> endobj 945 0 obj <> endobj 1325 0 obj <> endobj 1326 0 obj <> endobj 962 0 obj <> 24]>> endobj 1327 0 obj <> endobj 963 0 obj <> 26]>> endobj 1328 0 obj <> endobj 964 0 obj <> 28]>> endobj 1329 0 obj <> endobj 965 0 obj <> 30]>> endobj 1330 0 obj <> endobj 966 0 obj <> 32]>> endobj 967 0 obj <> 34]>> endobj 1331 0 obj <> endobj 1332 0 obj <> endobj 1333 0 obj <> endobj 1334 0 obj <> endobj 968 0 obj <> 40]>> endobj 1335 0 obj <> endobj 969 0 obj <> 43]>> endobj 1336 0 obj <> endobj 1337 0 obj <> endobj 1523 0 obj <>/P 1337 0 R/K 1>> endobj 1338 0 obj <> endobj 1339 0 obj <> endobj 1340 0 obj <> endobj 971 0 obj <>]>> endobj 972 0 obj <>]>> endobj 973 0 obj <>]>> endobj 974 0 obj <> 11<>]>> endobj 975 0 obj <>]>> endobj 976 0 obj <>]>> endobj 977 0 obj <>]>> endobj 978 0 obj <> 19<>]>> endobj 979 0 obj <>]>> endobj 980 0 obj <>]>> endobj 981 0 obj <>]>> endobj 982 0 obj <> 27<>]>> endobj 983 0 obj <>]>> endobj 984 0 obj <>]>> endobj 985 0 obj <>]>> endobj 1341 0 obj <> 990 0 R<> 991 0 R<> 992 0 R<>]/Type/StructElem/P 945 0 R/Pg 33 0 R/S/P>> endobj 986 0 obj <>]>> endobj 987 0 obj <><>]>> endobj 988 0 obj <>]>> endobj 1342 0 obj <> endobj 1343 0 obj <> endobj 990 0 obj <><>]>> endobj 991 0 obj <>]>> endobj 992 0 obj <><> 5]>> endobj 1344 0 obj <> endobj 993 0 obj <>]>> endobj 994 0 obj <><>]>> endobj 995 0 obj <> 12]>> endobj 996 0 obj <><>]>> endobj 997 0 obj <>]>> endobj 998 0 obj <>]>> endobj 999 0 obj <><>]>> endobj 1000 0 obj <>]>> endobj 1001 0 obj <>]>> endobj 1002 0 obj <><>]>> endobj 1003 0 obj <>]>> endobj 1004 0 obj <><>]>> endobj 1005 0 obj <>]>> endobj 1006 0 obj <> 34<>]>> endobj 1007 0 obj <>]>> endobj 1008 0 obj <>]>> endobj 1009 0 obj <> 40<>]>> endobj 1010 0 obj <> 42<>]>> endobj 1345 0 obj <> endobj 1011 0 obj <>]>> endobj 1346 0 obj <> endobj 1347 0 obj <> endobj 1348 0 obj <> endobj 1012 0 obj <>]>> endobj 1013 0 obj <><>]>> endobj 1349 0 obj <> endobj 1014 0 obj <>]>> endobj 1015 0 obj <> 57<>]>> endobj 1350 0 obj <> endobj 1351 0 obj <>]/Type/StructElem/P 945 0 R/Pg 66 0 R/S/P>> endobj 1016 0 obj <>]>> endobj 1017 0 obj <> 63<>]>> endobj 1353 0 obj <>/S/Table>> endobj 1352 0 obj <> endobj 1524 0 obj <> endobj 1526 0 obj <> endobj 1527 0 obj <>/P 1526 0 R/K 67>> endobj 1528 0 obj <>/P 1526 0 R/K 68>> endobj 1529 0 obj <>/P 1526 0 R/K 69>> endobj 1530 0 obj <>/P 1526 0 R/K 70>> endobj 1525 0 obj <> endobj 1531 0 obj <> endobj 1540 0 obj <> endobj 1541 0 obj <> endobj 1542 0 obj <> endobj 1543 0 obj <> endobj 1532 0 obj <> endobj 1544 0 obj <> endobj 1545 0 obj <> endobj 1546 0 obj <> endobj 1547 0 obj <> endobj 1533 0 obj <> endobj 1548 0 obj <> endobj 1549 0 obj <> endobj 1550 0 obj <> endobj 1551 0 obj <> endobj 1534 0 obj <> endobj 1552 0 obj <> endobj 1553 0 obj <> endobj 1554 0 obj <> endobj 1555 0 obj <> endobj 1535 0 obj <> endobj 1556 0 obj <> endobj 1557 0 obj <> endobj 1558 0 obj <> endobj 1559 0 obj <> endobj 1536 0 obj <> endobj 1560 0 obj <> endobj 1561 0 obj <> endobj 1562 0 obj <> endobj 1563 0 obj <> endobj 1537 0 obj <> endobj 1564 0 obj <> endobj 1565 0 obj <> endobj 1566 0 obj <> endobj 1567 0 obj <> endobj 1538 0 obj <> endobj 1568 0 obj <> endobj 1569 0 obj <> endobj 1570 0 obj <> endobj 1571 0 obj <> endobj 1539 0 obj <> endobj 1572 0 obj <> endobj 1573 0 obj <> endobj 1574 0 obj <> endobj 1575 0 obj <> endobj 1354 0 obj <> endobj 1355 0 obj <> 1023 0 R<> 1024 0 R<> 1025 0 R<>]/Type/StructElem/P 945 0 R/Pg 112 0 R/S/P>> endobj 1357 0 obj <>/S/Table>> endobj 1356 0 obj <> endobj 1576 0 obj <> endobj 1578 0 obj <> endobj 1579 0 obj <>/P 1578 0 R/K 5>> endobj 1580 0 obj <>/P 1578 0 R/K 6>> endobj 1581 0 obj <>/P 1578 0 R/K 7>> endobj 1582 0 obj <>/P 1578 0 R/K 8>> endobj 1583 0 obj <>/P 1578 0 R/K 9>> endobj 1584 0 obj <>/P 1578 0 R/K 10>> endobj 1585 0 obj <>/P 1578 0 R/K 11>> endobj 1586 0 obj <>/P 1578 0 R/K 12>> endobj 1587 0 obj <>/P 1578 0 R/K 13>> endobj 1588 0 obj <>/P 1578 0 R/K 14>> endobj 1577 0 obj <> endobj 1589 0 obj <> endobj 1600 0 obj <> endobj 1601 0 obj <> endobj 1602 0 obj <> endobj 1603 0 obj <> endobj 1604 0 obj <> endobj 1605 0 obj <> endobj 1606 0 obj <> endobj 1607 0 obj <> endobj 1608 0 obj <> endobj 1609 0 obj <> endobj 1590 0 obj <> endobj 1610 0 obj <> endobj 1611 0 obj <> endobj 1612 0 obj <> endobj 1613 0 obj <> endobj 1614 0 obj <> endobj 1615 0 obj <> endobj 1616 0 obj <> endobj 1617 0 obj <> endobj 1618 0 obj <> endobj 1619 0 obj <> endobj 1591 0 obj <> endobj 1620 0 obj <> endobj 1621 0 obj <> endobj 1622 0 obj <> endobj 1623 0 obj <> endobj 1624 0 obj <> endobj 1625 0 obj <> endobj 1626 0 obj <> endobj 1627 0 obj <> endobj 1628 0 obj <> endobj 1629 0 obj <> endobj 1592 0 obj <> endobj 1630 0 obj <> endobj 1631 0 obj <> endobj 1632 0 obj <> endobj 1633 0 obj <> endobj 1634 0 obj <> endobj 1635 0 obj <> endobj 1636 0 obj <> endobj 1637 0 obj <> endobj 1638 0 obj <> endobj 1639 0 obj <> endobj 1593 0 obj <> endobj 1640 0 obj <> endobj 1641 0 obj <> endobj 1642 0 obj <> endobj 1643 0 obj <> endobj 1644 0 obj <> endobj 1645 0 obj <> endobj 1646 0 obj <> endobj 1647 0 obj <> endobj 1648 0 obj <> endobj 1649 0 obj <> endobj 1594 0 obj <> endobj 1650 0 obj <> endobj 1651 0 obj <> endobj 1652 0 obj <> endobj 1653 0 obj <> endobj 1654 0 obj <> endobj 1655 0 obj <> endobj 1656 0 obj <> endobj 1657 0 obj <> endobj 1658 0 obj <> endobj 1659 0 obj <> endobj 1595 0 obj <> endobj 1660 0 obj <> endobj 1661 0 obj <> endobj 1662 0 obj <> endobj 1663 0 obj <> endobj 1664 0 obj <> endobj 1665 0 obj <> endobj 1666 0 obj <> endobj 1667 0 obj <> endobj 1668 0 obj <> endobj 1669 0 obj <> endobj 1596 0 obj <> endobj 1670 0 obj <> endobj 1671 0 obj <> endobj 1672 0 obj <> endobj 1673 0 obj <> endobj 1674 0 obj <> endobj 1675 0 obj <> endobj 1676 0 obj <> endobj 1677 0 obj <> endobj 1678 0 obj <> endobj 1679 0 obj <> endobj 1597 0 obj <> endobj 1680 0 obj <> endobj 1681 0 obj <> endobj 1682 0 obj <> endobj 1683 0 obj <> endobj 1684 0 obj <> endobj 1685 0 obj <> endobj 1686 0 obj <> endobj 1687 0 obj <> endobj 1688 0 obj <> endobj 1689 0 obj <> endobj 1598 0 obj <> endobj 1690 0 obj <> endobj 1691 0 obj <> endobj 1692 0 obj <> endobj 1693 0 obj <> endobj 1694 0 obj <> endobj 1695 0 obj <> endobj 1696 0 obj <> endobj 1697 0 obj <> endobj 1698 0 obj <> endobj 1699 0 obj <> endobj 1599 0 obj <> endobj 1700 0 obj <> endobj 1701 0 obj <> endobj 1702 0 obj <> endobj 1703 0 obj <> endobj 1704 0 obj <> endobj 1705 0 obj <> endobj 1706 0 obj <> endobj 1707 0 obj <> endobj 1708 0 obj <> endobj 1709 0 obj <> endobj 1358 0 obj <> endobj 1019 0 obj <>]>> endobj 1020 0 obj <>]>> endobj 1359 0 obj <>/P 945 0 R/K 131>> endobj 1360 0 obj <> endobj 1021 0 obj <><>]>> endobj 1023 0 obj <>]>> endobj 1024 0 obj <> 3<>]>> endobj 1025 0 obj <>]>> endobj 1361 0 obj <> endobj 1026 0 obj <>]>> endobj 1027 0 obj <><>]>> endobj 1028 0 obj <>]>> endobj 1362 0 obj <> endobj 1029 0 obj <>]>> endobj 1030 0 obj <><>]>> endobj 1363 0 obj <> endobj 1364 0 obj <> endobj 1031 0 obj <>]>> endobj 1365 0 obj <> endobj 1366 0 obj <> endobj 1032 0 obj <>]>> endobj 1033 0 obj <>]>> endobj 1034 0 obj <> 29<>]>> endobj 1035 0 obj <>]>> endobj 1036 0 obj <>]>> endobj 1037 0 obj <>]>> endobj 1038 0 obj <>]>> endobj 1039 0 obj <>]>> endobj 1040 0 obj <>]>> endobj 1041 0 obj <> 43<>]>> endobj 1042 0 obj <>]>> endobj 1367 0 obj <> endobj 1368 0 obj <> endobj 1043 0 obj <>]>> endobj 1044 0 obj <> 51]>> endobj 1369 0 obj <> 1050 0 R<>]/S/P>> endobj 1045 0 obj <> 54<>]>> endobj 1046 0 obj <>]>> endobj 1047 0 obj <>]>> endobj 1048 0 obj <><>]>> endobj 1050 0 obj <><>]>> endobj 1370 0 obj <> 1053 0 R<> 1054 0 R<> 1055 0 R<> 1056 0 R<>]/Type/StructElem/P 945 0 R/Pg 160 0 R/S/P>> endobj 1051 0 obj <>]>> endobj 1371 0 obj <>/P 945 0 R/K 7>> endobj 1372 0 obj <> endobj 1373 0 obj <>/P 945 0 R/K 9>> endobj 1374 0 obj <> endobj 1053 0 obj <><>]>> endobj 1054 0 obj <> 3]>> endobj 1055 0 obj <>]>> endobj 1056 0 obj <>]>> endobj 1375 0 obj <>]/Type/StructElem/P 945 0 R/Pg 168 0 R/S/P>> endobj 1377 0 obj <>/S/Table>> endobj 1376 0 obj <> endobj 1711 0 obj <> endobj 1713 0 obj <> endobj 1714 0 obj <>/P 1713 0 R/K 12>> endobj 1715 0 obj <>/P 1713 0 R/K 13>> endobj 1716 0 obj <>/P 1713 0 R/K 14>> endobj 1717 0 obj <>/P 1713 0 R/K 15>> endobj 1712 0 obj <> endobj 1718 0 obj <> endobj 1729 0 obj <> endobj 1730 0 obj <> endobj 1731 0 obj <> endobj 1732 0 obj <> endobj 1719 0 obj <> endobj 1733 0 obj <> endobj 1734 0 obj <> endobj 1735 0 obj <> endobj 1736 0 obj <> endobj 1720 0 obj <> endobj 1737 0 obj <> endobj 1738 0 obj <> endobj 1739 0 obj <> endobj 1740 0 obj <> endobj 1721 0 obj <> endobj 1741 0 obj <> endobj 1742 0 obj <> endobj 1743 0 obj <> endobj 1744 0 obj <> endobj 1722 0 obj <> endobj 1745 0 obj <> endobj 1746 0 obj <> endobj 1747 0 obj <> endobj 1748 0 obj <> endobj 1723 0 obj <> endobj 1749 0 obj <> endobj 1750 0 obj <> endobj 1751 0 obj <> endobj 1752 0 obj <> endobj 1724 0 obj <> endobj 1753 0 obj <> endobj 1754 0 obj <> endobj 1755 0 obj <> endobj 1756 0 obj <> endobj 1725 0 obj <> endobj 1757 0 obj <> endobj 1758 0 obj <> endobj 1759 0 obj <> endobj 1760 0 obj <> endobj 1726 0 obj <> endobj 1761 0 obj <> endobj 1762 0 obj <> endobj 1763 0 obj <> endobj 1764 0 obj <> endobj 1727 0 obj <> endobj 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<>]>> endobj 1106 0 obj <> 3<>]>> endobj 1107 0 obj <>]>> endobj 1108 0 obj <>]>> endobj 1387 0 obj <> endobj 1388 0 obj <> endobj 1389 0 obj <> endobj 1390 0 obj <> endobj 1109 0 obj <>]>> endobj 1110 0 obj <> 15<>]>> endobj 1391 0 obj <> endobj 1392 0 obj <>]/Type/StructElem/P 945 0 R/Pg 239 0 R/S/P>> endobj 1111 0 obj <>]>> endobj 1112 0 obj <><>]>> endobj 1394 0 obj <>/S/Table>> endobj 1393 0 obj <> endobj 1835 0 obj <> endobj 1837 0 obj <> endobj 1838 0 obj <>/P 1837 0 R/K 25>> endobj 1839 0 obj <>/P 1837 0 R/K 26>> endobj 1840 0 obj <>/P 1837 0 R/K 27>> endobj 1836 0 obj <> endobj 1841 0 obj <> endobj 1857 0 obj <> endobj 1858 0 obj <> endobj 1859 0 obj <> endobj 1842 0 obj <> endobj 1860 0 obj <> endobj 1861 0 obj <> endobj 1862 0 obj <> endobj 1843 0 obj <> endobj 1863 0 obj <> endobj 1864 0 obj <> endobj 1865 0 obj <> endobj 1844 0 obj <> endobj 1866 0 obj <> endobj 1867 0 obj <> endobj 1868 0 obj <> endobj 1845 0 obj <> endobj 1869 0 obj <> endobj 1870 0 obj <> endobj 1871 0 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